Santosh has completed his PhD from Center for Cellular and Molecular Biology (CCMB), Hyderabad, India under the supervision of Dr. Rajan Sankaranarayanan and which was funded by DST-INSPIRE (Dept. of Science and Technology, India) fellowship. His Ph.D. thesis was mainly focused in deciphering the “Mechanism and Physiological Role of Chiral Proofreading Modules Involved in Translation of the Genetic Code”. In the process of his Ph.D. training, he has acquired expertise in the field of structural and molecular cell biology. After a brief postdoc in India, Santosh moved to Germany to join the Dikic group to pursue the structural and physiological consequences of phosphoribosyl-ubiquitination. He move on in 2023.